Dan is a Visiting Scholar at Duke University's Marine Lab, and in 2018 will join Louisiana State University's Department of Oceanographic and Coastal Sciences as an Assistant Professor. His research focuses on the connectivity of coral reef metapopulations, and on the resilience of marine metapopulations to perturbations and climate changes. He has continued interest in mesophotic coral ecosystems as potential refugia from climate change - and how these refuges and adjacent habitats interact through the exchange of individuals or genes to produce persistent metapopulations. Tools employed by Dan's lab include simulation and ecological modeling, coral reproductive histology, and deep genome sequencing; all aimed at deciphering the mechanisms that control reef population connectivity, and how connectivity influences persistence in time.

  Please note that only publications relevant to mesophotic reefs are indexed.

	Fertile fathoms: Deep reproductive refugia for threatened shallow corals | article Holstein DM, Smith TB, Gyory J, Paris CB (2015) Sci Rep 5:12407
	Modeling vertical coral connectivity and mesophotic refugia | article Holstein DM, Paris CB, Vaz AC, Smith TB (2016) Coral Reefs 35:23-37
	Depth-independent reproduction in the reef coral Porites astreoides from shallow to mesophotic zones | article Holstein DM, Smith TB, Paris CB (2016) PLoS ONE 11:e0146068
	Vertical connectivity in mesophotic coral ecosystems | thesis Holstein DM (2013)
	Potential structuring forces on a shelf edge upper Mesophotic Coral Ecosystem in the US Virgin Islands | article Smith TB, Brandtneris VW, Canals M, Brandt ME, Martens J, Brewer RS, Kadison E, Kammann M, Keller J, Holstein DM (2016) Front Mar Sci 3:115
	Growth rates of Porites astreoides and Orbicella franksi in mesophotic habitats surrounding St. Thomas, US Virgin Islands | article Groves SH, Holstein DM, Enochs IC, Kolodzeij G, Manzello DP, Brandt ME, Smith TB (2018) Coral Reefs 37: 345–354
	The United States Virgin Islands | chapter Smith TB, Brandt ME, Brandtneris VW, Ennis RS, Groves SH, Habtes S, Holstein DM, Kadison E, Nemeth RS (2019) in: Mesophotic Coral Ecosystems (Springer) by Loya Y, Puglise KA, Bridge TCL
	Disturbance in Mesophotic Coral Ecosystems and Linkages to Conservation and Management | chapter Smith TB, Holstein DM, Ennis RS (2019) in: Mesophotic Coral Ecosystems () by Loya Y, Puglise KA, Bridge TCL
	Mesophotic coral refuges following multiple disturbances | article Bloomberg J, Holstein DM (2021) Coral Reefs 40:821-834
	Predicting the distribution of threatened orbicellid corals in shallow and mesophotic reef ecosystems | article Egan KE, Viehman TS, Holstein DM, Poti M, Groves SH, Smith TB (2021) Mar Ecol Prog Ser 667:61-81
	Ecosystem Services of Mesophotic Coral Ecosystems and a Call for Better Accounting | chapter Holstein DM, Fletcher P, Groves SH, Smith TB (2019) in: Mesophotic Coral Ecosystems (Springer) by Loya Y, Puglise KA, Bridge TCL

  SCUBA (open-circuit or unspecified)
  Rebreather
  Coring / drilling
  Snorkeling